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Florida State University
Florida Resources and
Environmental Analysis Center
at the Institute of Science
and Public Affairs

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Natural Communities Descriptions

This summary presents the hierarchical classification and brief descriptions of 82 Natural Communities developed by Florida Natural Areas Inventory and identified as collectively constituting the original, natural biological associations of Florida.

A Natural Community is defined as a distinct and recurring assemblage of populations of plants, animals, fungi and microorganisms naturally associated with each other and their physical environment. For more complete descriptions, see the Guide to the Natural Communities of Florida.

The levels of the hierarchy are:

Natural Community Category - defined by hydrology and vegetation.

Natural Community Groups - defined by landform, substrate, and vegetation.

Natural Community Type - defined by landform and substrate; soil moisture condition; climate; fire; and characteristic vegetation.



TERRESTRIAL - Upland habitats dominated by plants which are not adapted to anaerobic soil conditions imposed by saturation or inundation for more than 10% of the growing season.

XERIC UPLANDS - very dry, deep, well-drained hills of sand with xeric-adapted vegetation.

Sandhill - upland with deep sand substrate; xeric; temperate; frequent fire (2-5 years); longleaf pine and/or turkey oak with wiregrass understory.

Scrub - old dune with deep fine sand substrate; xeric; temperate or subtropical; occasional or rare fire (20 - 80 years); sand pine and/or scrub oaks and/or rosemary and lichens.

Xeric Hammock - upland with deep sand substrate; xeric-mesic; temperate or subtropical; rare or no fire; live oak and/or sand live oak and/or laurel oak and/or other oaks, sparkleberry, saw palmetto.

 

COASTAL UPLANDS - substrate and vegetation influenced primarily by such coastal (maritime) processes as erosion, deposition, salt spray, and storms.

Beach Dune - active coastal dune with sand substrate; xeric; temperate or subtropical; occasional or rare fire; sea oats and/or mixed salt-spray tolerant grasses and herbs.

Coastal Berm - old bar or storm debris with sand/shell substrate; xeric-mesic; subtropical or temperate; rare or no fire; buttonwood, mangroves, and/or mixed halophytic herbs and/or shrubs and trees.

Coastal Grassland - coastal flatland with sand substrate; xeric-mesic; subtropical or temperate; occasional fire; grasses, herbs, and shrubs with or without slash pine and/or cabbage palm.

Coastal Rock Barren - flatland with exposed limestone substrate; xeric; subtropical; no fire; algae, mixed halophytic herbs and grasses, and/or cacti and stunted shrubs and trees.

Coastal Strand - stabilized coastal dune with sand substrate; xeric; subtropical or temperate; occasional or rare fire; dense saw palmetto and/or seagrape and/or mixed stunted shrubs, yucca, and cacti.

Maritime Hammock - stabilized coastal dune with sand substrate; xeric-mesic; subtropical or temperate; rare or no fire; mixed hardwoods and/or live oak.

Shell Mound - Indian midden with shell substrate; xeric-mesic; subtropical or temperate; rare or no fire; mixed hardwoods.

 

MESIC UPLANDS - dry to moist hills of sand with varying amounts of clay, silt or organic material; diverse mixture of broadleaved and needleleaved temperate woody species.

Slope Forest - steep slope on bluff or in sheltered ravine; sand/clay substrate; mesic-hydric; temperate; rare or no fire; magnolia, beech, spruce pine, Shumard oak, Florida maple, mixed hardwoods.

Upland Glade - upland with calcareous rock and/or clay substrate; hydric-xeric; temperate; sparse mixed grasses and herbs with occasional stunted trees and shrubs, e.g., eastern red cedar.

Upland Hardwood Forest - upland with sand/clay and/or calcareous substrate; mesic; temperate; rare or no fire; spruce pine, magnolia, beech, pignut hickory, white oak, and mixed hardwoods.

Upland Mixed Forest - upland with sand/clay substrate; mesic; temperate; rare or no fire; loblolly pine and/or shortleaf pine and/or laurel oak and/or magnolia and spruce pine and/or mixed hardwoods.

Upland Pine Forest - upland with sand/clay substrate; mesic-xeric; temperate; frequent or occasional fire; longleaf pine and/or loblolly pine and/or shortleaf pine, southern red oak, wiregrass.

 

ROCKLANDS - low, generally flat limestone outcrops with tropical vegetation; or limestone exposed through karst activities with tropical or temperate vegetation.

Pine Rockland - flatland with exposed limestone substrate; mesic-xeric; subtropical; frequent fire; south Florida slash pine, palms and/or hardwoods, and mixed grasses and herbs.

Rockland Hammock - flatland with limestone substrate; mesic; subtropical; rare or no fire; mixed tropical hardwoods, often with live oak.

Sinkhole - karst feature with steep limestone walls; mesic-hydric; subtropical or temperate; no fire; ferns, herbs, shrubs, and hardwoods.

 

MESIC FLATLANDS - flat, moderately well-drained sandy substrates with admixture of organic material, often with a hard pan.

Dry Prairie - flatland with sand substrate; mesic-xeric; subtropical or temperate; annual or frequent fire; wiregrass, saw palmetto, and mixed grasses and herbs.

Mesic Flatwoods - flatland with sand substrate; mesic; subtropical or temperate; frequent fire; slash pine and/or longleaf pine with saw palmetto, gallberry and/or wiregrass or cutthroat grass understory.

Mesic Hammock - flatland with sand substrate; mesic; temperate; occasional or rare fire; live oak, cabbage palm, and saw palmetto.

Prairie Hammock - flatland with sand/organic soil over marl or limestone substrate; mesic; subtropical; occasional or rare fire; live oak and/or cabbage palm.

Scrubby Flatwoods - flatland with sand substrate; xeric-mesic; subtropical or temperate; occasional fire; longleaf pine or slash pine with scrub oaks and wiregrass understory.


PALUSTRINE - Wetlands dominated by plants adapted to anaerobic substrate conditions imposed by substrate saturation or inundation during 10% or more of the growing season. Includes non-tidal wetlands; tidal wetlands with ocean derived salinities less than 0.5 ppt and dominance by salt-intolerant species; small (less than 8 ha), shallow (less than 2 m deep at low water) water bodies without wave-formed or bedrock shoreline; and inland brackish or saline wetlands.

WET FLATLANDS - flat, poorly drained sand, marl or limestone substrates.

Hydric Hammock - lowland with sand/clay/organic soil, often over limestone; mesic-hydric; subtropical or temperate; rare or no fire; water oak, cabbage palm, red cedar, red maple, bays, hackberry, hornbeam, blackgum, needle palm, and mixed hardwoods.

Marl Prairie - flatland with marl over limestone substrate; seasonally inundated; tropical; frequent to no fire; sawgrass, spikerush, and/or mixed grasses, sometimes with dwarf cypress.

Wet Flatwoods - flatland with sand substrate; seasonally inundated; subtropical or temperate; frequent fire; vegetation characterized by slash pine or pond pine and/or cabbage palm with mixed grasses and herbs.

Wet Prairie - flatland with sand substrate; seasonally inundated; subtropical or temperate; annual or frequent fire; maidencane, beakrush, spikerush, wiregrass, pitcher plants, St. John's wort, mixed herbs.

 

SEEPAGE WETLANDS - sloped or flat sands or peat with high moisture levels maintained by downslope seepage; wetland and mesic woody and/or herbaceous vegetation.

Baygall - wetland with peat substrate at base of slope; maintained by downslope seepage, usually saturated and occasionally inundated; subtropical or temperate; rare or no fire; bays and/or dahoon holly and/or red maple and/or mixed hardwoods.

Seepage Slope - wetland on or at base of slope with organic/sand substrate; maintained by downslope seepage, usually saturated but rarely inundated; subtropical or temperate; frequent or occasional fire; sphagnum moss, mixed grasses and herbs or mixed hydrophytic shrubs.

 

FLOODPLAIN WETLANDS - flat, alluvial sand or peat substrates associated with flowing water courses and subjected to flooding but not permanent inundation; wetland or mesic woody and herbaceous vegetation.

Bottomland Forest - flatland with sand/clay/organic substrate; occasionally inundated; temperate; rare or no fire; water oak, red maple, beech, magnolia, tuliptree, sweetgum, bays, cabbage palm, and mixed hardwoods.

Floodplain Forest - floodplain with alluvial substrate of sand, silt, clay or organic soil; seasonally inundated; temperate; rare or no fire; diamondleaf oak, overcup oak, water oak, swamp chestnut oak, blue palmetto, cane, and mixed hardwoods.

Floodplain Marsh - floodplain with organic/sand/alluvial substrate; seasonally inundated; subtropical; frequent or occasional fire; maidencane, pickerelweed, sagittaria spp., buttonbush, and mixed emergents.

Floodplain Swamp - floodplain with organic/alluvial substrate; usually inundated; subtropical or temperate; rare or no fire; vegetation characterized by cypress, tupelo, black gum, and/or pop ash.

Freshwater Tidal Swamp - river mouth wetland, organic soil with extensive root mat; inundated with freshwater in response to tidal cycles; rare or no fire; cypress, bays, cabbage palm, gums and/or cedars.

Slough - broad, shallow channel with peat over mineral substrate; seasonally inundated, flowing water; subtropical; occasional or rare fire; pop ash and/or pond apple or water lily.

Strand Swamp - broad, shallow channel with peat over mineral substrate; seasonally inundated, flowing water; subtropical; occasional or rare fire; cypress and/or willow.

Swale - broad, shallow channel with sand/peat substrate; seasonally inundated, flowing water; subtropical or temperate; frequent or occasional fire; sawgrass, maidencane, pickerelweed, and/or mixed emergents.

 

BASIN WETLANDS - shallow, closed basin with outlet usually only in time of high water; peat or sand substrate, usually inundated; wetland woody and/or herbaceous vegetation.

Basin Marsh - large basin with peat substrate; seasonally inundated; temperate or subtropical; frequent fire; sawgrass and/or cattail and/or buttonbush and/or mixed emergents.

Basin Swamp - large basin with peat substrate; seasonally inundated, still water; subtropical or temperate; occasional or rare fire; vegetation characterized by cypress, blackgum, bays and/or mixed hardwoods.

Bog - wetland on deep peat substrate; moisture held by sphagnum mosses, soil usually saturated, occasionally inundated; subtropical or temperate; rare fire; sphagnum moss and titi and/or bays and/or dahoon holly, and/or mixed hydrophytic shrubs.

Coastal Interdunal Swale - long narrow depression wetlands in sand/peat-sand substrate; seasonally inundated, fresh to brackish, still water; temperate; rare fire; graminoids and mixed wetland forbs.

Depression Marsh - small rounded depression in sand substrate with peat accumulating toward center; seasonally inundated, still water; subtropical or temperate; frequent or occasional fire; maidencane, fire flag, pickerelweed, and mixed emergents, may be in concentric bands.

Dome Swamp - rounded depression in sand/limestone substrate with peat accumulating toward center; seasonally inundated, still water; subtropical or temperate; occasional or rare fire; cypress, blackgum, or bays, often tallest in center.


LACUSTRINE - Non-flowing wetlands of natural depressions lacking persistent emergent vegetation except around the perimeter.

Clastic Upland Lake - generally irregular basin in clay uplands; predominantly with inflows, frequently without surface outflow; clay or organic substrate; colored, acidic, soft water with low mineral content (sodium, chloride, sulfate); oligo-mesotrophic to eutrophic.

Coastal Dune Lake - basin or lagoon influenced by recent coastal processes; predominantly sand substrate with some organic matter; salinity variable among and within lakes, and subject to saltwater intrusion and storm surges; slightly acidic, hard water with high mineral content (sodium, chloride).

Coastal Rockland Lake - shallow basin influence by recent coastal processes; predominantly barren oolitic or Miami limestone substrate; salinity variable among and within lakes, and subject to saltwater intrusion, storm surges and evaporation (because of shallowness); slightly alkaline, hard water with high mineral content (sodium, chloride).

Flatwoods/Prairie Lake - generally shallow basin in flatlands with high water table; frequently with a broad littoral zone; still water or flow-through; sand or peat substrate; variable water chemistry, but characteristically colored to clear, acidic to slightly alkaline, soft to moderately hard water with moderate mineral content (sodium, chloride, sulfate); oligo-mesotrophic to eutrophic.

Marsh lake - generally shallow, open water area within wide expanses of freshwater marsh; still water or flow-through; peat, sand or clay substrate; occurs in most physiographic regions; variable water chemistry, but characteristically highly colored, acidic, soft water with moderate mineral content (sodium, chloride, sulfate); oligo-mesotrophic to eutrophic.

River Floodplain Lake - meander scar, backwater, or larger flow-through body within major river floodplains; sand, alluvial or organic substrate; colored, alkaline or slightly acidic, hard or moderately hard water with high mineral content (sulfate, sodium, chloride, calcium, magnesium); mesotrophic to eutrophic.

Sandhill Upland Lake - generally rounded solution depression in deep sandy uplands or sandy uplands shallowly underlain by limestone; predominantly without surface inflows/outflows; typically sand substrate with organic accumulations toward middle; clear, acidic moderately soft water with varying mineral content; ultra-oligotrophic to mesotrophic.

Sinkhole Lake - typically deep, funnel-shaped depression in limestone base; occurs in most physiographic regions; predominantly without surface inflows/outflows, but frequently with connection to the aquifer; clear, alkaline, hard water with high mineral content (calcium, bicarbonate, magnesium).

Swamp Lake - generally shallow, open water area within basin swamps; still water or flow-through; peat, sand or clay substrate; occurs in most physiographic regions; variable water chemistry, but characteristically highly colored, acidic, soft water with moderate mineral content (sodium, chloride, sulfate); oligo-mesotrophic to eutrophic.


RIVERINE - Natural, flowing waters from their source to the downstream limits of tidal influence and bounded by channel banks.

Alluvial Stream - lower perennial or intermittent/seasonal watercourse characterized by turbid water with suspended silt, clay, sand and small gravel; generally with a distinct, sediment-derived (alluvial) floodplain and a sandy, elevated natural levee just inland from the bank.

Blackwater Stream - perennial or intermittent/seasonal watercourse characterized by tea-colored water with a high content of particulate and dissolved organic matter derived from drainage through swamps and marshes; generally lacking an alluvial floodplain.

Seepage Stream - upper perennial or intermittent/seasonal watercourse characterized by clear to lightly colored water derived from shallow groundwater seepage.

Spring-run Stream - perennial watercourse with deep aquifer headwaters and characterized by clear water, circumneutral pH and, frequently, a solid limestone bottom.


SUBTERRANEAN - Twilight, middle and deep zones of natural chambers overlain by the earth's crust and characterized by climatic stability and assemblages of trogloxenic, troglophilic, and troglobitic organisms.

Aquatic Cave - cavernicolous area permanently or periodically submerged; often characterized by troglobitic crustaceans and salamanders; includes high energy systems which receive large quantities of organic detritus and low energy systems.

Terrestrial Cave - cavernicolous area lacking standing water; often characterized by bats, such as Myotis spp., and other terrestrial vertebrates and invertebrates; includes interstitial areas above standing water such as fissures in the ceiling of caves.


MARINE/ESTUARINE (The distinction between the Marine and Estuarine Natural Communities is often subtle, and the natural communities types found under these two community categories have the same descriptions. For these reasons they have been grouped together.) - Subtidal, intertidal and supratidal zones of the sea, landward to the point at which seawater becomes significantly diluted with freshwater inflow from the land.

Consolidated Substrate - expansive subtidal, intertidal and supratidal area composed primarily of nonliving compacted or coherent and relatively hard, naturally formed mass of mineral matter (e.g., coquina limerock and relic reefs); octocorals, sponges, stony corals, nondrift macrophytic algae, blue-green mat-forming algae and seagrasses sparse, if present.

Unconsolidated Substrate - expansive subtidal, intertidal and supratidal area composed primarily of loose mineral matter (e.g., coralgal, gravel, marl, mud, sand and shell); octocorals, sponges, stony corals, nondrift macrophytic algae, blue-green mat-forming algae and seagrasses sparse, if present.

Octocoral Bed - expansive subtidal area occupied primarily by living sessile organisms of the Class Anthozoa, Subclass Octocorallia (e.g., soft corals, horny corals, sea fans, sea whips, and sea pens); sponges, stony corals, nondrift macrophytic algae and seagrasses spares, if present.

Sponge Bed - expansive subtidal area occupied primarily by living sessile organisms of the Phylum Porifera (e.g., sheepswool sponge, Florida loggerhead sponge and branching candle sponge); octocorals, stony corals, nondrift macrophytic algae and seagrasses sparse, if present.

Coral Reef - expansive subtidal area with elevational gradient or relief and occupied primarily by living sessile organisms of the Class Hydrozoa (e.g., fire corals and hydrocorals) and Class Anthozoa, Subclass Zoantharia (e.g., stony corals and black corals); includes deepwater bank reefs, fringing barrier reefs, outer bank reefs and patch reefs, some of which may contain distinct zones of assorted macrophytes, octocorals, & sponges.

Mollusk Reef - substantial subtidal or intertidal area with relief from concentrations of sessile organisms of the Phylum Mollusca, Class Bivalvia (e.g., molluscs, oysters, & worm shells); octocorals, sponges, stony corals, macrophytic algae and seagrasses sparse, if present.

Worm Reef - substantial subtidal or intertidal area with relief from concentrations of sessile, tubicolous organisms of the Phylum Annelida, Class Polychaeta (e.g., chaetopterids and sabellarids); octocorals, sponges, stony corals, macrophytic algae and seagrasses sparse, if present.

Algal Bed - expansive subtidal, intertidal or supratidal area, occupied primarily by attached thallophytic or mat-forming prokaryotic algae (e.g, halimeda, blue-green algae); octocorals, sponges, stony corals and seagrasses sparse, if present.

Grass Bed - expansive subtidal or intertidal area, occupied primarily by rooted vascular macrophytes, (e.g., shoal grass, halophila, widgeon grass, manatee grass and turtle grass); may include various epiphytes and epifauna; octocorals, sponges, stony corals, and attached macrophytic algae sparse, if present.

Composite Substrate - expansive subtidal, intertidal, or supratidal area, occupied primarily by Natural Community elements from more than one Natural Community category (e.g., Grass Bed and Algal Bed species; Octocoral and Algal Bed species); includes both patchy and evenly distributed occurrences.

Tidal Marsh - expansive intertidal or supratidal area occupied primarily by rooted, emergent vascular macrophytes (e.g., cord grass, needlerush, saw grass, saltwort, saltgrass and glasswort); may include various epiphytes and epifauna.

Tidal Swamp - expansive intertidal and supratidal area occupied primarily by woody vascular macrophytes (e.g., black mangrove, buttonwood, red mangrove, and white mangrove); may include various epiphytes and epifauna.


DEFINITIONS OF TERMS Terrestrial and Palustrine Natural Communities

Physiography
Upland - high area in region with significant topographic relief; generally undulating
Lowland - low area in region with or without significant topographic relief; generally flat to gently sloping
Flatland - generally level area in region without significant topographic relief; flat to gently sloping
Basin - large, relatively level lowland with slopes confined to the perimeter or isolated interior locations
Depression - small depression with sloping sides, deepest in center and progressively shallower towards the perimeter
Floodplain - lowland adjacent to a stream; topography influenced by recent fluvial processes
Bottomland - lowland not on active floodplain; sand/clay/organic substrate

Hydrology
occasionally inundated - surface water present only after heavy rains and/or during flood stages
seasonally inundated - surface water present during wet season and flood periods
usually inundated - surface water present except during droughts

Climatic Affinity of the Flora
tropical
- community generally occurs in practically frost-free areas
subtropical - community generally occurs in areas that experience occasional frost, but where freezing temperatures are not frequent enough to cause true winter dormancy
temperate - community generally occurs in areas that freeze often enough that vegetation goes into winter dormancy

Fire
annual fire - burns about every 1-2 years
frequent fire - burns about every 3-7 years
occasional fire - burns about every 8-25 years
rare fire - burns about every 26-100 years
no fire - community develops only when site goes more than 100 years without burning


LATIN NAMES OF PLANTS MENTIONED IN NATURAL COMMUNITY DESCRIPTIONS
anise - Illicium floridanum
bays:
     swamp bay -Persea palustris
     gordonia -Gordonia lasianthus
     sweetbay -Magnolia virgiana
beakrush - Rhynchospora spp.
beech - Fagus grandifolia
blackgum - Nyssa biflora
blue palmetto - Sabal minor
bluestem - Andropogon spp.
buttonbush - Cephalanthus occidentalis
cabbage palm - Sabal palmetto
cacti - Opuntia and Harrisia spp.,
     predominantly stricta and pentagonus
cane - Arundinaria gigantea or A. tecta
cattail - Typha spp.
cedars:
     red cedar - Juniperus silicicola
     white cedar - Chamaecyparis thyoides or C. henryi
cladonia - Cladonia spp.
cypress - Taxodium distichum
dahoon holly - Ilex cassine
diamondleaf oak - Quercus laurifolia
fire flag - Thalia geniculata
Florida maple - Acer barbatum
gallberry - Ilex glabra
gums:
     tupelo - Nyssa aquatica
     blackgum - Nyssa biflora
     Ogeechee gum - Nyssa ogeche
hackberry - Celtis laevigata
hornbeam - Carpinus caroliniana
laurel oak - Quercus hemisphaerica
live oak - Quercus virginiana
loblolly pine - Pinus taeda
longleaf pine - Pinus palustris
magnolia - Magnolia grandiflora
maidencane - Panicum hemitomon
needle palm - Rhapidophyllum hystrix
overcup oak - Quercus lyrata
pickerel weed - Pontederia cordata or P. lanceolata
pignut hickory - Carya glabra
pop ash - Fraxinus caroliniana
pond apple - Annona glabra
pond pine - Pinus serotina
pyramid magnolia - Magnolia pyramidata
railroad vine - Ipomoea pes-caprae
red cedar - Juniperus silicicola
red maple - Acer rubrum
red oak - Quercus falcata
rosemary - Ceratiola ericoides
sagittaria - Sagittaria lancifolia
sand pine - Pinus clausa
saw palmetto - Serenoa repens
sawgrass - Cladium jamaicensis
scrub oaks - Quercus geminata, Q. chapmanii, Q. myrtifolia,Q. inopina
sea oats - Uniola paniculata
seagrape - Coccoloba uvifera
shortleaf pine - Pinus echinata
Shumard oak - Quercus shumardii
slash pine - Pinus elliottii
sphagnum moss - Sphagnum spp.
spikerush - Eleocharis spp.
spruce pine - Pinus glabra
St. John's wort - Hypericum spp.
swamp chestnut oak - Quercus prinus
sweetgum - Liquidambar styraciflua
titi - Cyrilla racemiflora, and Cliftonia monophylla
tuliptree - Liriodendron tulipfera
tupelo - Nyssa aquatica
turkey oak - Quercus laevis
water oak - Quercus nigra
waterlily - Nymphaea odorata
white cedar - Chamaecyparis thyoides
white oak - Quercus alba
willow - Salix caroliniana
yucca - Yucca aloifolia